He was born in 1931 in Miyazaki Prefecture, on Kyūshū
Island, Japan. He was associate professor and professor of biology at Brown
University from 1969 to 1972 and professor of population genetics at the Center for
Demographic and Population Genetics, University of Texas at Houston, from 1972
He is a theoretical population geneticist and evolutionary biologist. Acting alone or working with his students, he has continuously developed new statistical theories of molecular evolution taking into account frontier knowledge of molecular biology. He has also developed several new concepts of evolutionary theory and proposed the theory of mutation-driven evolution.
قدیمیترین باغ گیاهشناسی ایران در شرف تخریب «به حکم استانداری»